New paper on cycadophytes from Madygen

Moisan, P., S. Voigt, C. Pott, M. Buchwitz, J. Schneider, and H. Kerp. in press. Cycadalean and bennettitalean foliage from the Triassic Madygen Lagerstätte (SW Kyrgyzstan, Central Asia). Review of Palaeobotany and Palynology. [DOI:10.1016/j.revpalbo.2010.11.008]

Philippe Moisan who is doing his Ph.D. in Münster (with paleobotanist Hans Kerp as his supervisor) studies the flora of the Triassic Madygen Fm. In his first paper on that issue he introduces cycadophyte finds collected between 2005 and 2009.

Many of the studied the specimen come from the same succession and locality as Madygenerpeton (there is also a small sketch of the sedimentary profile, see Fig. 2).

One thing I learned from this study was that so-called "xeromorphic features", i.e. plant features that are usually the consequence of an adaptation to aridity, cannot only occur in xerophytes, i.e. in plants adapted to dry environments, but (for other reasons) in hygrophytic and halophytic plants as well.

Indications for aridity, such as desiccation crack horizons or or seasonally drying-out ponds and rivers or wide-spread red bed sediments are lacking in Madygen. Thus, according to Philippe's interpretation, "xeromorphism" in Madygen plants probably served other purposes than the xeromorphism of xerophytes (e.g. "self-cleaning of the leaf surface, regulation of excessive radiation and leaf temperature, mechanical defense against phytophagous insects").

Three recent papers on chroniosuchians

Buchwitz M, Voigt S. 2010. Peculiar carapace structure of a Triassic chroniosuchian implies evolutionary shift in trunk flexibility. Journal of Vertebrate Paleontology 30: 1697-1708. [Link]

Schoch RR, Voigt S, Buchwitz M. 2010. A chroniosuchid from the Triassic of Kyrgyzstan and analysis of chroniosuchian relationships. Zoological Journal of the Linnaean Society 160: 515-530. [Link]

Klembara J, Clack J, Čerňanský A. 2010. The anatomy of palate of Chroniosaurus dongusensis (Chroniosuchia, Chroniosuchidae) from the Upper Permian of Russia. Palaeontology 53: 1147-1153. [Link]

The redescription of the Chroniosaurus dongusensis palate by Klembara and colleagues adds further data to the morphological dataset provided by Clack and Klembara (2009) in their revision of C. dongusensis on the basis of a new specimen (which is the most complete of any yet known chroniosuchian). According to the updated phylogenetic analysis from the 2010 paper Chroniosaurus as the only included chroniosuchian taxon formed the sister group of embolomeres.

Schoch and colleagues (me included) describe Madygenerpeton pustulatus, a new species of chroniosuchians from the Middle to Late Triassic of Central Asia with a highly derived skull morphology and a carapace that was chroniosuchid-like in many aspects. The find shows that one lineage of chroniosuchids survived the Permian-Triassic boundary (by 20 or so million years).

The authors discuss characteristics uniting chroniosuchians with "higher reptiliomorphs" and unlike the approach of Klembara and colleagues their cladistic analysis, which includes five chroniosuchian taxa, results in a position of chroniosuchians somewhat closer to amniotes than to embolomeres. Chroniosaurus comes out as the closest relative of Madygenerpeton (both share the characteristic ornamentation of the skull and osteoderms besides other features).

Buchwitz & Voigt consider the functionality of chroniosuchian carapaces, comparing them to archosaur osteoderm systems. They argue that chroniosuchian carapaces basically served terrestrial locomotion but that the higher lateral flexibility of the Madygenerpeton osteoderm system was linked to a secondary increase in undulation swimming capability.

Reference:
Clack JA, Klembara J. 2009. An articulated specimen of Chroniosaurus dongusensis, and the morphology and relationships of the chroniosuchids. Special Papers in Palaeontology 81: 15-42. [Link]

Palges Meeting October 2010 in Munich

The 80 th Annual Meeting of the German Paleontological Society took place from the 6th through the 8th October 2010 within the halls of the Bavarian State Collection for Geology and Paleontology in Munich.

Of particular interest for me was the session on Early Mesozoic vertebrates chaired by the Rauhut couple and Richard Butler as it united many interesting characters, such as Silvio Renesto, Martin Ezcurra, Rainer Schoch, and Daniela Schwarz-Wings and covered a variety of Triassic vertebrates including archosaurs, temnospondyls, and bony fish.

With 5 talks and 6 posters our small Freibergian working group had quite a number of contributions this year (my prof Jörg Schneider was talking about Paleozoic cockroaches from China, Olaf Elicki about Cambrian trace fossils from Africa and the Middle East, Frederik Spindler about the evolution of haptodonts and other early synapsids, Jan Fischer about oxygen isotope signals in Permian and Triassic freshwater shark teeth and I had a talk on osteoderm histology and the Chroniosuchia). My colleagues Ilja Kogan and Jan Fischer won the 1st poster prize with their poster entitled "The Madygen lake deposits: A unique multi-taxa kindergarten for Triassic fisches" - which is quite an achievement as normally the winner comes from the host institute of the Palges Meeting.

The image on the right shows me in front of a poster entitled "Paleontology in the German Wikipedia" [pdf].

Even though there are many private collectors and paleontology enthusisasts in Germany, Austria, and Switzerland you won't find much about "regional paleontology" in the German Wikipedia which was the reason for my colleagues and me to introduce some aspects of Wikipedian (Pop-)Sciencewriting.

The poster praised the advantages Wikipedia can have if it is reasonably incorporated in public outreach campaigns and we commented critically on the dinosaur focus which increases the already biased public image of what paleontology is about.

Madygenerpeton pustulatus: first description finally out

Schoch, R. R., S. Voigt, and M. Buchwitz. 2010. A chroniosuchid from the Triassic of Kyrgyzstan and analysis of chroniosuchian relationships. Zoological Journal of the Linnean Society 160(3): 515-530. [Abstract]

Madygen trace fossil paper

Voigt, S. and D. Hoppe. 2010. Mass Occurrence of Penetrative Trace Fossils in Triassic Lake Deposits (Kyrgyzstan, Central Asia). Ichnos 17:1-11. [Link]

Besides the exquisite soft body preservation of insects and tetrapods within some parts of the lacustrine succession, the Triassic Madygen lake shows a rich inventary of invertebrate trace fossils, studied by my colleague from Freiberg Sebastian Voigt.

The interesting point about these ichnofossil assemblages is that they demonstrate a certain differentiation of the lake ground in better and less well aerated zones, displaying different degrees of bioturbation and abundances of indicative ichnotaxa.

You can imagine that fresh water lake grounds only became inhabited stepwise after the conquest of land by animals, so these trace fossil assemblages mark a certain evolutionary level of lake ecosystems, otherwise rarely documented in detail from the Middle to Late Triassic.